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New contributions to the taxonomic revision of the ctenostylid flies (Diptera: Tephritoidea: Ctenostylidae) Valery A. Korneyev I.I.Schmalhausen Institute of Zoology, Kiev, Ukraine This presentation does not consist a publication in the sense of the Code of Zoological Nomenclature and all the new names are used only provisionally !
What are ctenostylids? ,[object Object]
History ,[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object],(C) To Chan (Hong Kong) (C) To Chan (Hong Kong)
[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object]
[object Object],[object Object],[object Object],[object Object],[object Object]
Taxonomic position ,[object Object],[object Object],[object Object]
Are ctenostylids Tephritoidea? ( pros  and  contras )
contras ,[object Object],[object Object],[object Object]
pros ,[object Object],[object Object],[object Object]
Morphology ,[object Object],Peristomal cavity large, anterior part of head strongly oblique (in profile), mainly consisting of modified peristomal cavity  (~as high as face in  Furciseta & Ramuliseta  or  2x as high as face in  Lochmostylia ,  Tauroscypson  and  Ctenostylum )  Furciseta  male Ramuliseta  female Lochmostylia  male Lochmostylia  female
Why holoptic eyes? In distantly related  Furciseta, Nepaliseta  and  Ctenostylum . ,[object Object],[object Object],[object Object],[object Object]
Antenna Furciseta  sp. (from Korneyev, 2010) Tauroscypson andinum  (from Aczel, 1956) Nepaliseta mirabilis  (from Korneyev, 2001) Female: arista:  (1)  with two branches;  (2)  with more than two branches;  (3)  disk-like.
Antenna M ale: arista :   simple, moderately long pubescent . Furciseta  sp. (from Korneyev, 2010) Tauroscypson andinum  (from Aczel, 1956) Nepaliseta mirabilis  (from Korneyev, 2001) Female  needs very sensitive mechanoreceptors  to feel air movements? that males do not need.  To look for the host? Furciseta plaumanni  (from Korneyev, 2010) Ramuliseta thaica  (from Korneyev, 2001)
Why membranous face? Lochmostylia lopesi  female Lochmostylia borgmeieri  female
Thorax Scutellum :  (1)  with two  setae ;  (2)  with more than two  setae ;  (3)  with brush of setulae, but no setae . Furciseta  sp.  Ctenostylum  sp. female Nepaliseta  sp. strongly widened, short, as long as head; proepisternum entire, transverse, notopleural triangle shortened, with 1-2 p npl seta(-e) . Swarming ?
Why short phallus? Ramuliseta thaica  male Lochmostylia borgmeieri  male Furciseta plaumanni  male Swarming ? Copulation on the flight?
Why no aculeus? Pyrgotomyia (Pyrgotini)   Africa Teretrura  (Teretrurinae) Chile Furciseta plaumanni  male Epice  (Toxurini) New Guinea
Phylogenetic Relationships ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Phylogenetic Relationships ,[object Object],[object Object],[object Object],[object Object],[object Object],[object Object],[object Object]
Parsymony analysis
New material accumulated  by Doug Yanega (L.A., California) and myself.  A new approach to the taxonomic revision  of the ctenostylids is needed.
2. Genus  Ramuliseta  Keiser, 1951 ,[object Object],[object Object]
A brief survey of known ctenostylid species ,[object Object],[object Object],[object Object]
3. Clade  Sinolochmostylia + Nepaliseta + R. thaica  group R2+3 and R4+5 strongly divergent. Female with short round fl.1 & disk-like arista. (not known for  “Ramuliseta ” spp.) ,[object Object],[object Object],[object Object],[object Object],[object Object],1 2 4 5 3
4.   Ramuliseta madagascariensis  group ,[object Object],[object Object],1 2
5.  Lochmostylia  Hendel, 1934 +  Tauroscypson  Curran, 1934 +  Ctenostylum  Macquart, 1851  ,[object Object],[object Object],[object Object],1 1 2 3
5.  Lochmostylia  Hendel, 1934 +  Tauroscypson  Curran, 1934 +  Ctenostylum  Macquart, 1851  ,[object Object],[object Object],1 2 2 2 1
5.  Lochmostylia  Hendel, 1934 +  Tauroscypson  Curran, 1934 +  Ctenostylum  Macquart, 1851  ,[object Object]
Thank you for attention!   6.  gen .  sp.

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V korneyev new contributions to the taxonomic revision s

  • 1. New contributions to the taxonomic revision of the ctenostylid flies (Diptera: Tephritoidea: Ctenostylidae) Valery A. Korneyev I.I.Schmalhausen Institute of Zoology, Kiev, Ukraine This presentation does not consist a publication in the sense of the Code of Zoological Nomenclature and all the new names are used only provisionally !
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  • 10.
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  • 13. Are ctenostylids Tephritoidea? ( pros and contras )
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  • 16.
  • 17.
  • 18. Antenna Furciseta sp. (from Korneyev, 2010) Tauroscypson andinum (from Aczel, 1956) Nepaliseta mirabilis (from Korneyev, 2001) Female: arista: (1) with two branches; (2) with more than two branches; (3) disk-like.
  • 19. Antenna M ale: arista : simple, moderately long pubescent . Furciseta sp. (from Korneyev, 2010) Tauroscypson andinum (from Aczel, 1956) Nepaliseta mirabilis (from Korneyev, 2001) Female needs very sensitive mechanoreceptors to feel air movements? that males do not need. To look for the host? Furciseta plaumanni (from Korneyev, 2010) Ramuliseta thaica (from Korneyev, 2001)
  • 20. Why membranous face? Lochmostylia lopesi female Lochmostylia borgmeieri female
  • 21. Thorax Scutellum : (1) with two setae ; (2) with more than two setae ; (3) with brush of setulae, but no setae . Furciseta sp. Ctenostylum sp. female Nepaliseta sp. strongly widened, short, as long as head; proepisternum entire, transverse, notopleural triangle shortened, with 1-2 p npl seta(-e) . Swarming ?
  • 22. Why short phallus? Ramuliseta thaica male Lochmostylia borgmeieri male Furciseta plaumanni male Swarming ? Copulation on the flight?
  • 23. Why no aculeus? Pyrgotomyia (Pyrgotini) Africa Teretrura (Teretrurinae) Chile Furciseta plaumanni male Epice (Toxurini) New Guinea
  • 24.
  • 25.
  • 27. New material accumulated by Doug Yanega (L.A., California) and myself. A new approach to the taxonomic revision of the ctenostylids is needed.
  • 28.
  • 29.
  • 30.
  • 31.
  • 32.
  • 33.
  • 34.
  • 35. Thank you for attention! 6. gen . sp.