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   Stage 1: Abiotic Synthesis of Organic
    Monomers
       Amino Acids
         Chains of nucleotides
           Chains of DNA bases
           Chains of RNA bases
         Building blocks of protein
     Sugars
     Lipids
     ATP
   Stage 2: Abiotic Synthesis of Polymers
       Monomers, such as amino acids, spontaneously
        fused together to form proteins.
   Stage 3: Origin of Self-Replicating Molecules

                    inheritance




Ribozyme: catalytic RNA used to fuel RNA replication
   Stage 4: Formation of
    Pre-cells
       Molecular packages with
        some properties of life.
       The gap between pre-cells
        and true cells is
        enormous!
         Natural Selection
The origin of
   eukaryotic cells
Endosymbiotic Theory

 Membrane bound nuclear
  material
 Organelles
 More complex than prokaryotic
  cells
 Ancestors to fungi, plants and
  animals
Concept Map
Section 17-2
                                      Evolution of Life
                           Early Earth was hot; atmosphere contained poisonous gases.

                                       Earth cooled and oceans condensed.

                           Simple organic molecules may have formed in the oceans..

                            Small sequences of RNA may have formed and replicated.

               First prokaryotes may have formed when RNA or DNA was enclosed in microspheres.

                          Later prokaryotes were photosynthetic and produced oxygen.

                      An oxygenated atmosphere capped by the ozone layer protected Earth.

                          First eukaryotes may have been communities of prokaryotes.

                                        Multicellular eukaryotes evolved.

                      Sexual reproduction increased genetic variability, hastening evolution.
Evolution = “Descent with Modification”
    1. ------------due to being spread over
       different habitats and therefore adapting
       and modifying themselves to fit the
       habitat.
    2. Natural Selection
        • A population of organisms can
          change over generations if certain
          inheritable traits leave more
          offspring than others and those
          others get the chance to reproduce
          and continue the lineage.
        • Evolutionary Adaptation
Greek Philosophers

• Plato’s Idealism

• Aristotle’s scala naturae (scale of
  nature)

 special creation of each species
 organisms were created in their current
  form
 the earth was only a few thousand years
  old
THOMAS MALTHUS

        economist & clergyman

        published
         An Essay on the Principle of
                Population



 Populations had an inherent tendency to increase
geometrically, while the resources needed to support this growth
increase slowly or not at all.

 Because of the continued growth of a species would outstrip
needed resources, growth would be limited.
JEAN-BAPTISTE de LAMARCK

                           French biologist
                           proposed that modern species
                            descended from other species
                           Lamarckism based on two theories:


1.   Inheritance of Acquired Characteristics
     - traits acquired by an individual during its life are passed to its
     offspring
2. Use and Disuse
     - organs of the body that were used extensively to cope with the
     environment became larger and stronger, while organs that
              were not used deteriorated
THE ROAD TO EVOLUTIONARY THEORY
Example:           Evolution Before Darwin
The Evolution of the Giraffe

Giraffes obtained their long necks from
previous giraffes who stretched to eat the
leaves of high tree branches.
Stretching increased the length of their
necks, and this acquired characteristic
was passed to the next generation.
Lamarckism

   first to present a unified theory that attempted to explain the
    changes in organisms from one generation to the next

   Although, mechanisms proposed for change was wrong, since
    acquired characteristics are not heritable!!!

       origin of species from preexisting species
       ability of organisms to adapt
GEORGES CUVIER
   French anatomist and
    naturalist and writer
   palaeontologist
   strongly opposed the
    concept of evolution
 history of living organisms recorded in layers of rock
   containing a succession of fossil species in chronological
   order

 fossils were organisms that had died in a series of
   catastrophes, after which extinct plants and animals were
   replaced by the immigration of distant species to the
   devastated region  Catastrophism
CHARLES DARWIN
   voyage on the H.M.S.
    Beagle (1836)
   published (1859)
    The Origin of Species
   first person who proposed
    a mechanistic approach to
    evolutionary thought
   the father of synthetic
    evolution
 Galapagos tortoise
 Darwin’s finches
 Most organisms on the islands
were endemic.
   The birds were very similar (common ancestry).

   They all adapted to different Niches.
       Niche = Your job in nature.
         What you eat
         Where you live (on ground, in a tree)


   Reproductive Isolation resulted in speciation.
Fossils chronologically
ordered in rock layers
Geographic distribution of species: Australian marsupials
HOMOLOGY
Notice that a
Chimp is more
genetically
related to a
human than to
an Old World
Monkey!
 species not specially created in their
  present forms, but had evolved from
  ancestral species
 proposed a mechanism for evolution:
            Natural Selection

   A population of organisms can
   change over time as a result of
   individuals with certain heritable
   characteristics leaving more offspring
   than other individuals.
Darwinism in Historical Context
GREGOR MENDEL
   Austrian biologist
   discovered the basic principles of
    heredity
   father of Classical Genetics


 Individual characteristics determined
   by inherited factors transmitted from
   parent to offspring.
HUGO                CARL               ERICH VON
     DE VRIES            CORRENS             TSCHERMAK


   rediscovered Mendel’s laws of heredity
   the start of rediscovering evolution in terms of Mendel’s
    ideas
JAMES WATSON
FRANCIS CRICK
   elucidated the structure of
    DNA (genetic material)
   DNA contains coded
    information which acts as a
    blueprint for the transfer of
    hereditary information from
    generation to generation
   mutation as the raw
    material for evolution
   A population is the smallest unit of evolution.
     Natural selection acts on individuals.
     However the evolutionary impact of natural selection is only
      apparent in tracking how a population changes over time.
   Population Genetics emphasizes the extensive genetic variation
    within populations and tracks the genetic make-up of populations
    over time.
     Not all variation in a population is inheritable.
     Only the genetic component of variation is relevant to natural
      selection.
     Many variable traits in a population result from the combined effect of
      several genes.
Genetic equilibrium- allele frequencies
remain constant.
 Populations should be in a state of equilibrium
  assuming:
   1. Large Population
   2. Random Mating
   3. No Migration
   4. No Mutation
   5. No natural selection occurs.
   Perfectly adapted organsim.
       Sharks
       Alligator
       Horseshoe Crab
   Stable envrionment.
       Water (Oceans)
   Speciation occurs relatively quickly with long
    periods of equilibrium in between.
       Speciation = The formation of a new species due to
        evolution.
   Evolution can occur without natural selection.
   Genetic Drift is a change in the gene pool due
    to a random event.
       Example- Mount Saint Helens Eruption.
   If the event is catastrophic and all the
    organisms are killed off = Mass extinction.
   = The formation of a new species due to evolution.
   Reproductive Isolation- Can result in speciation. It
    is the separation of members of a species due to
    some barrier.
     Islands
     Mountains
     Roads
     Oil Pipes
•Speciation occurs only
with the evolution of
reproductive barriers
between the isolated
population and its parent
population.
•If speciation occurs
during geographic
separation, the new
species will not breed
with its ancestral
population, even if the
two populations should
come back into contact.
   Allopatric Speciation
     A population forms a
      new species while
      geographically isolated
      from its parent
      population.
   Sympatric Speciation
     A small population
      becomes a new species
      in the midst of a parent
      population
   Does not widely occur among
    animals but may account for
    over 25% of all plant species.
Notice how the hybrid
 bread wheat has a set
 of chromosomes from
 each of its ancestors:
 T. monococcum (AA),
 Triticum (BB), T.
 turgidum (AA BB), T.
 tauschii (DD)


What can you notice about
T. aestivum that might
make it a good hybrid and
the most important wheat
species today?
   Convergent Evolution- Organisms of different
    ancestry adapting to a similar environment.
       Produces homologous Structures.
       Example: bats and Butterflies wings.
         Dolphin
         Penguin
         Shark



   Divergent Evolution- Organisms of common
    ancestry adapting to diverse environments.
       Produces analogous Structures.
       Example: Human Arm and Bird wing.
1. Genetic Drift
  Def: A change in the gene pool of a small population due to chance.

   The best measure of Darwinian fitness is the number of fertile
   offspring an individual leaves.
2. Gene Flow
   Def: The genetic exchange with another population.

3. Mutations
      A change in an organism’s DNA sequence.
      Ultimate source of genetic variability.

4. Natural Selection
       Directional Selection (selecting in favor of an extreme phenotype)
       Disruptive (Diversifying) Selection (leads to a balance between two or
        more contrasting phenotypic forms)
       Stabilizing Selection (maintains variation in a narrow range)
•Resistant Genes

    •Immediate
    Benefits
    •Long term
    Disaster
    •Evolution direct
    connection to
    daily lives
   Def: Major biological changes
    evident in fossil record.
   CONTRAST:
    MICROEVOLUTION
   Speciation
     Nonbranching evolution
       (transform a population
       enough for it to be designated
       a new species.)
     Branching evolution (splits a
       lineage into two or more
       species)
   Ernst Mayr
       Studied the diversity of birds in New Guinea (1927)
       Biological species concept
          Species = “groups of interbreeding natural populations that are
            reproductively isolated from other such groups.”
          Reproductive isolation blocks exchange of genes between species and
            keeps their gene pools separate.

• Reproductive barriers between species
  • Zygote: fertilized egg
  • Pre-zygotic (factors that impede
  mating between species or hinder
  fertilization of eggs if mating is
  attempted)
  • Post-zygotic (mechanisms that
  operate should interspecies mating
  actually occur and form hybrid
  zygotes)
Isolating Mechanisms


          Courtship




       Sterile
Spurts of
Slow          relatively
adaptations   rapid
              change
Flowchart
          Section 17-4


                                                         Species


                                                          that are



                       Unrelated                                                                Related



        form                           in        under                    under        in                      in

                                  Similar                    Intense
Inter-relationshiops                                                                Small                   Different
                               environments               environmental
                                                                                  populations             environments
                                                             pressure

    can undergo                    can undergo             can undergo            can undergo             can undergo



    Coevolution                    Convergent                                     Punctuated                Adaptive
                                                            Extinction
                                    evolution                                     equilibrium               radiation
   Systematics
       Reconstructing evolutionary history
       Radioactive dating = tool
       The study of biological diversity: past and present
   Taxonomy
       Identification, naming and classification of species
       Pioneer: Carolus Linnaeus (1707-1778)
         Binomial Nomenclature
         Homo sapien
Hierarchical
   Classification


  Which step is more
      specific?
    Class or Genus?



   How is Domain
Eukarya different from
Bacteria and Archaea?
•Evolutionary
history of a
species.
•Tree is based on
homologous
structures, NOT
analogous.
•Two species will
have more
common
nucleotide
sequences based
on how recently
they branched
from their
common ancestor.
The Computer
Age
Archaea-
Eubacteria bacteria
                      ?

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Bio 40s evolution

  • 1. Stage 1: Abiotic Synthesis of Organic Monomers  Amino Acids  Chains of nucleotides  Chains of DNA bases  Chains of RNA bases  Building blocks of protein  Sugars  Lipids  ATP
  • 2. Stage 2: Abiotic Synthesis of Polymers  Monomers, such as amino acids, spontaneously fused together to form proteins.
  • 3. Stage 3: Origin of Self-Replicating Molecules inheritance Ribozyme: catalytic RNA used to fuel RNA replication
  • 4. Stage 4: Formation of Pre-cells  Molecular packages with some properties of life.  The gap between pre-cells and true cells is enormous!  Natural Selection
  • 5. The origin of eukaryotic cells Endosymbiotic Theory  Membrane bound nuclear material  Organelles  More complex than prokaryotic cells  Ancestors to fungi, plants and animals
  • 6. Concept Map Section 17-2 Evolution of Life Early Earth was hot; atmosphere contained poisonous gases. Earth cooled and oceans condensed. Simple organic molecules may have formed in the oceans.. Small sequences of RNA may have formed and replicated. First prokaryotes may have formed when RNA or DNA was enclosed in microspheres. Later prokaryotes were photosynthetic and produced oxygen. An oxygenated atmosphere capped by the ozone layer protected Earth. First eukaryotes may have been communities of prokaryotes. Multicellular eukaryotes evolved. Sexual reproduction increased genetic variability, hastening evolution.
  • 7.
  • 8. Evolution = “Descent with Modification” 1. ------------due to being spread over different habitats and therefore adapting and modifying themselves to fit the habitat. 2. Natural Selection • A population of organisms can change over generations if certain inheritable traits leave more offspring than others and those others get the chance to reproduce and continue the lineage. • Evolutionary Adaptation
  • 9. Greek Philosophers • Plato’s Idealism • Aristotle’s scala naturae (scale of nature)  special creation of each species  organisms were created in their current form  the earth was only a few thousand years old
  • 10. THOMAS MALTHUS  economist & clergyman  published An Essay on the Principle of Population  Populations had an inherent tendency to increase geometrically, while the resources needed to support this growth increase slowly or not at all.  Because of the continued growth of a species would outstrip needed resources, growth would be limited.
  • 11. JEAN-BAPTISTE de LAMARCK  French biologist  proposed that modern species descended from other species  Lamarckism based on two theories: 1. Inheritance of Acquired Characteristics - traits acquired by an individual during its life are passed to its offspring 2. Use and Disuse - organs of the body that were used extensively to cope with the environment became larger and stronger, while organs that were not used deteriorated
  • 12. THE ROAD TO EVOLUTIONARY THEORY Example: Evolution Before Darwin The Evolution of the Giraffe Giraffes obtained their long necks from previous giraffes who stretched to eat the leaves of high tree branches. Stretching increased the length of their necks, and this acquired characteristic was passed to the next generation.
  • 13. Lamarckism  first to present a unified theory that attempted to explain the changes in organisms from one generation to the next  Although, mechanisms proposed for change was wrong, since acquired characteristics are not heritable!!!  origin of species from preexisting species  ability of organisms to adapt
  • 14. GEORGES CUVIER  French anatomist and naturalist and writer  palaeontologist  strongly opposed the concept of evolution  history of living organisms recorded in layers of rock containing a succession of fossil species in chronological order  fossils were organisms that had died in a series of catastrophes, after which extinct plants and animals were replaced by the immigration of distant species to the devastated region  Catastrophism
  • 15. CHARLES DARWIN  voyage on the H.M.S. Beagle (1836)  published (1859) The Origin of Species  first person who proposed a mechanistic approach to evolutionary thought  the father of synthetic evolution
  • 16.
  • 17.  Galapagos tortoise Darwin’s finches Most organisms on the islands were endemic.
  • 18.
  • 19. The birds were very similar (common ancestry).  They all adapted to different Niches.  Niche = Your job in nature.  What you eat  Where you live (on ground, in a tree)  Reproductive Isolation resulted in speciation.
  • 20.
  • 22. Geographic distribution of species: Australian marsupials
  • 24. Notice that a Chimp is more genetically related to a human than to an Old World Monkey!
  • 25.  species not specially created in their present forms, but had evolved from ancestral species  proposed a mechanism for evolution: Natural Selection A population of organisms can change over time as a result of individuals with certain heritable characteristics leaving more offspring than other individuals.
  • 26.
  • 28. GREGOR MENDEL  Austrian biologist  discovered the basic principles of heredity  father of Classical Genetics  Individual characteristics determined by inherited factors transmitted from parent to offspring.
  • 29. HUGO CARL ERICH VON DE VRIES CORRENS TSCHERMAK  rediscovered Mendel’s laws of heredity  the start of rediscovering evolution in terms of Mendel’s ideas
  • 30. JAMES WATSON FRANCIS CRICK  elucidated the structure of DNA (genetic material)  DNA contains coded information which acts as a blueprint for the transfer of hereditary information from generation to generation  mutation as the raw material for evolution
  • 31. A population is the smallest unit of evolution.  Natural selection acts on individuals.  However the evolutionary impact of natural selection is only apparent in tracking how a population changes over time.  Population Genetics emphasizes the extensive genetic variation within populations and tracks the genetic make-up of populations over time.  Not all variation in a population is inheritable.  Only the genetic component of variation is relevant to natural selection.  Many variable traits in a population result from the combined effect of several genes.
  • 32. Genetic equilibrium- allele frequencies remain constant.  Populations should be in a state of equilibrium assuming:  1. Large Population  2. Random Mating  3. No Migration  4. No Mutation  5. No natural selection occurs.
  • 33.
  • 34. Perfectly adapted organsim.  Sharks  Alligator  Horseshoe Crab  Stable envrionment.  Water (Oceans)
  • 35. Speciation occurs relatively quickly with long periods of equilibrium in between.  Speciation = The formation of a new species due to evolution.
  • 36.
  • 37. Evolution can occur without natural selection.  Genetic Drift is a change in the gene pool due to a random event.  Example- Mount Saint Helens Eruption.  If the event is catastrophic and all the organisms are killed off = Mass extinction.
  • 38. = The formation of a new species due to evolution.  Reproductive Isolation- Can result in speciation. It is the separation of members of a species due to some barrier.  Islands  Mountains  Roads  Oil Pipes
  • 39. •Speciation occurs only with the evolution of reproductive barriers between the isolated population and its parent population. •If speciation occurs during geographic separation, the new species will not breed with its ancestral population, even if the two populations should come back into contact.
  • 40. Allopatric Speciation  A population forms a new species while geographically isolated from its parent population.  Sympatric Speciation  A small population becomes a new species in the midst of a parent population
  • 41. Does not widely occur among animals but may account for over 25% of all plant species.
  • 42. Notice how the hybrid bread wheat has a set of chromosomes from each of its ancestors: T. monococcum (AA), Triticum (BB), T. turgidum (AA BB), T. tauschii (DD) What can you notice about T. aestivum that might make it a good hybrid and the most important wheat species today?
  • 43. Convergent Evolution- Organisms of different ancestry adapting to a similar environment.  Produces homologous Structures.  Example: bats and Butterflies wings.  Dolphin  Penguin  Shark  Divergent Evolution- Organisms of common ancestry adapting to diverse environments.  Produces analogous Structures.  Example: Human Arm and Bird wing.
  • 44. 1. Genetic Drift Def: A change in the gene pool of a small population due to chance. The best measure of Darwinian fitness is the number of fertile offspring an individual leaves.
  • 45. 2. Gene Flow Def: The genetic exchange with another population. 3. Mutations  A change in an organism’s DNA sequence.  Ultimate source of genetic variability. 4. Natural Selection  Directional Selection (selecting in favor of an extreme phenotype)  Disruptive (Diversifying) Selection (leads to a balance between two or more contrasting phenotypic forms)  Stabilizing Selection (maintains variation in a narrow range)
  • 46. •Resistant Genes •Immediate Benefits •Long term Disaster •Evolution direct connection to daily lives
  • 47. Def: Major biological changes evident in fossil record.  CONTRAST: MICROEVOLUTION  Speciation  Nonbranching evolution (transform a population enough for it to be designated a new species.)  Branching evolution (splits a lineage into two or more species)
  • 48. Ernst Mayr  Studied the diversity of birds in New Guinea (1927)  Biological species concept  Species = “groups of interbreeding natural populations that are reproductively isolated from other such groups.”  Reproductive isolation blocks exchange of genes between species and keeps their gene pools separate. • Reproductive barriers between species • Zygote: fertilized egg • Pre-zygotic (factors that impede mating between species or hinder fertilization of eggs if mating is attempted) • Post-zygotic (mechanisms that operate should interspecies mating actually occur and form hybrid zygotes)
  • 49. Isolating Mechanisms Courtship Sterile
  • 50. Spurts of Slow relatively adaptations rapid change
  • 51. Flowchart Section 17-4 Species that are Unrelated Related form in under under in in Similar Intense Inter-relationshiops Small Different environments environmental populations environments pressure can undergo can undergo can undergo can undergo can undergo Coevolution Convergent Punctuated Adaptive Extinction evolution equilibrium radiation
  • 52. Systematics  Reconstructing evolutionary history  Radioactive dating = tool  The study of biological diversity: past and present  Taxonomy  Identification, naming and classification of species  Pioneer: Carolus Linnaeus (1707-1778)  Binomial Nomenclature  Homo sapien
  • 53. Hierarchical Classification Which step is more specific? Class or Genus? How is Domain Eukarya different from Bacteria and Archaea?
  • 54. •Evolutionary history of a species. •Tree is based on homologous structures, NOT analogous. •Two species will have more common nucleotide sequences based on how recently they branched from their common ancestor.