JJ Thomson discovered the electron through his work with cathode ray tubes in the late 1800s. This led him to propose the plum pudding model of the atom, with negatively charged electrons embedded in a positively charged medium. Robert Millikan later directly measured the charge of individual electrons through his oil drop experiment in 1905, enabling him to calculate the electron's mass. Ernest Rutherford then used the gold foil experiment to discover the nucleus as the small, dense, positively charged center of atoms through deflections of alpha particles in 1910.